Word: knowleses
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Knowles, who has been a faculty member at Harvard since 1974, has published more than 250 research papers and is widely considered one of the best enzymologists in the world.
After his appointment as Dean of the Faculty in 1991, Knowles withdrew from research to focus on administrative duties. His last graduate student received her Ph.D. in 1994.
By replacing Professor Thorn, Knowles willmaintain the current ratio of scientists tonon-scientists among the HHMI trustees.
"One trustee dedicated to research in themedical and life sciences is being replaced bysomebody at least with his roots in the researchand teaching in the life sciences," Knowles noted.
Such academic qualifications, according toJarmul, were more important in deciding on asuccessor to Thorn than Knowles' affiliation withHarvard.